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The water vascular system is a modified part of coelom & consists of a system of sea water filled canals having certain corpuscles.
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Introduction :- The water vascular system is a modified part of coelom & consists of a system of sea water filled canals having certain corpuscles. It plays most vital role in the locomotion of the animals & comprises madreporite stone canal, ring canal, radial canal, Tiedman's body, lateral canals & tube feet. (1) Madriporite :- The madreporite is a rounded calcareous plate occurring on the aboral surface of the central disc in inter-radial position. Its surface bears a number of radiating, narrow, straight or wavy grooves or furrows. Each furrow contains many minute pores at its bottom. Each pore leads into a very short, fine, tubular pore-canal. Which passes inward in the substance of the madreporite. There may be about 200 pores and pore-canal. The pore-canals unite to form the collecting canals. Which open into an ampulla beneath the madreporite. Fig : Water vascular system of Starfish (2) Stone Canal :- The ampulla opens into a "S" shaped stone canal. The stone canal extends downwards (orally) and opens into a ring canal, around the mouth. The walls of stone canal are supported by a series of calcareous ringd. The lumen of stone canal is lined by very tall flagellated cells. in embryonic
stages and young Asterias, the stone canal remains a simple tube but in adult Asterias, lumen of stone canal possesses a prominent ridge with two spirally rolled lamellae. Fig : Diagram of Starfish (3) Ring Canal :- The Ring canal or water ring is located to the inner side of the peristomial ring of ossicles and directly above (aboral) to the hyponeural ring sinus. It is wide and pentagonal or five sided. Fig : Star fish
Fig : Function of water vascular system of Star fish (1) Locomotion :- The water vascular system is used mainly for locomotion. The inner wall of the water vascular canals are provided with cilia. The beating of the cilia causes the seawater to enter through the madreporite. Finally, the seawater reaches the tube feet and their ampullae. The ampullae contract ; the valves at the junction of the lateral canals and tube feet, prevent the flow of water into radial canals. The water is forced into the podia. The podia are elongated and protected out through the ambulacral groove. Then the suckers are applied to the substratum. The tube feet now contract & push the body forward. The water from the tube feet is pushed into the ampulla. Hence, the tube feet shorten. The suckers are released. Then the ampulla contracts & the whole process is repeated. (2) Food Capture :- The tube feet are used to capture the prey. The suckers are used to open the shells of molluscas. (3) Attachment :- The Starfish can be attached to the rocks by the tube feet.